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FIGURE 1 Ubiquitin–Proteasome System: A Complexed but Coordinated System The UPS is a highly controlled mechanism of protein degradation and turnover in cells, starting with monomeric ubiquitin, a small protein with a molecular mass of approximately 8 kDa ( The Proteasome as a Target for Cancer Therapy The 20S proteasome is a 700-kDa large complex formed by four stacked rings, two outer α-rings and two inner β-rings